Plant phenological responses to global warming are well studied. However, while many locations are experiencing increased temperatures, some locations are experiencing climate cooling. Little work has been conducted to understand plant phenological responses to cooling trends, much less the combined effects of cooling and other factors, such as changing precipitation. Furthermore, studies based on herbarium specimens have been instrumental in demonstrating plant responses to global warming; but to our knowledge, herbarium records have not been used to investigate responses to cooling.
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