This Field Guide to Conservation in Cities captures the collective experience of The Nature Conservancy and Center for Whole Communities (CWC) from 2014 to 2016, as they initiated a network of urban conservation programs in 12 cities across the United States. This North America Region program, generously supported by The JPB Foundation, was designed to expand The Conservancy’s work in cities. Its objectives were to culturally and operationally prepare them to engage in urban environmental issues in and with low-income communities, using an ongoing learning network to develop and scale successful conservation programs in urban areas.